By Christina Rexrode And Robin Sidel 

Banks just can't figure out what to do with their branches.

Firms across the country are tossing out their tellers, adding salespeople and remaking their drive-through lanes. The moves are the latest cost-cutting experiments for the industry as customers move in droves to online and mobile banking.

The efforts are proving tricky given that banks want customers to come to branches for moneymaking financial products such as loans or credit cards, but also have to serve the millions of Americans who still regularly make deposits and withdraw cash at the teller line.

When Capital One Financial Corp.'s new Boston office opens this summer, it won't have any tellers at all.

The firm is expanding a "cafe" format in which visitors can get gourmet coffee and free Wi-Fi but can't sit with a banker who will open an account for them or drop off a loan application.

Instead, employees steer customers to its website and answer questions about the bank's services. As it has in other locations, the company also will likely host occasional alcohol-free "appy hours" that promote the smartphone apps of local businesses.

Capital One sees the cafe format, which it is using at several locations around the country, including some already operating in Boston, as a way to promote the bank's brand without employing large numbers of tellers, a spokeswoman said.

Bank of America Corp. soon will begin converting about 9,000 tellers to "relationship bankers" who can direct customers to high-tech ATMs or show them how to deposit a check via smartphone. J.P. Morgan Chase & Co., PNC Financial Services Group Inc. and other banks are making similar moves.

U.S. banks operated 94,725 branches last year, the lowest number since 2005, according to the Federal Deposit Insurance Corp.

Despite banks cutting costs sharply in recent years, as many as one-third of those branches remain unprofitable, says consulting firm Simon-Kucher & Partners.

New ATMs that dispense exact change or that connect customers to a remote teller via video are meant to guide customers away from the teller for simple transactions. For banks, it is cheaper when a customer deposits a check at the ATM, and cheaper still when they deposit by taking a picture of it with a smartphone.

But converting branches into sales centers will be no easy task.

"Most banks don't have a clear vision of where to take the branch," says Robert Meara, senior analyst in the banking group at consulting firm Celent.

Only about 5% of customers applied for a new account or product on their most recent visit to a bank branch, according to market-research firm J.D. Power, and some estimates put the proportion even lower.

"Nobody buys a home equity line of credit on impulse," said Steven Reider, president of Bancography, which advises banks on their branches.

In some cases, the changes risk turning off customers.

Grace Raymond was frustrated after she stopped by a Bank of America in Ocala, Fla., to pay off her mortgage and encountered a "lobby full of customers" and no one to help clarify whether she needed to go to a financial adviser or a teller. After a long wait for a financial adviser, Ms. Raymond learned that she could have just gone to a teller, she said.

At a happy hour with friends later "I was so angry and annoyed that I couldn't calm down," said Ms. Raymond, who is 79 years old. "And normally I'm a pretty nice person."

At Bank of America's shareholder meeting in May, a shareholder asked CEO Brian Moynihan about long lines at some branches due to fewer tellers. Mr. Moynihan said the bank is trying to keep up with changing customer behavior.

A handful of banks are closing drive-through lanes or trying ways to make them more efficient.

Wells Fargo & Co. recently tested a new process in which customers go online to receive a one-time pass code to speed up the authentication process at the drive-through. The bank is analyzing the results.

PNC has toyed with "order ahead" ideas, which might allow customers to, for example, say ahead of time that they are going to withdraw $100 from the ATM, then scan their phone or enter a pass code once they get to the machine.

The bank says it is treading carefully.

"One thing we hear from customers: 'If you change the inside, fine. But don't mess up the drive-through,'" said Todd Barnhart, head of retail distribution at PNC.

Some firms are simply shuttering branches. Fifth Third Bancorp last month announced that it will close or sell 100 branches, or 8% of its offices. Like many of its peers, the Cincinnati-based bank also is expanding its use of employees who can handle a variety of functions.

At a Capital One cafe in New York recently, Jennifer Oliver used the ATM located inside the building and then walked to the back for a cup of coffee from the Peet's Coffee & Tea counter. The physician assistant said she was pleased to get a 50% discount on coffee by paying with her Capital One debit card, but said she was unlikely to become a regular since she rarely carries cash and has only visited a branch twice in the past eight years.

"I feel badly for all the bank tellers who don't have jobs anymore," she said.

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